William Mobley, PhD, received a PhD in neuro-biobehavioral sciences (1974) and an MD (1976) and completed an internship in pathology and residency in pediatrics (1979) at Stanford University. He was resident and fellow in neurology at Johns Hopkins University (1979-1982). He completed postdoctoral training at Walter Reed Institute for Research in Washington D.C. before joining the Department of Neurology at the University of California, San Francisco (1985). In 1997, he moved to Stanford as Chair of Neurology and then as Neurosciences Institute Director (2006). At the University of California, San Diego, he served as Chair of Neurosciences (2009-2017) and now as Associate Dean for Neurosciences Initiatives (2017-). He is a member of the National Academy of Medicine (2004) and fellow of the American Association for the Advancement of Sciences (2006). Using model systems, his research focuses on the molecular bases of neurotrophic factor signaling, mechanisms of neurodegeneration, and the impact of neurodegeneration on trophic signaling.